Glossary of pelagic biogeography r. K. Johnson†, B. J. Zahuranec*, D. Boltovskoy and A. C. Pierrot-Bults



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particle spectrum

espectro de partículas

Distribution of biomass in different size categories determined by the diameter of a sphere equivalent in volume (v) to the original particle multiplied by the number of particles (n).


patchiness

agregación

The tendency of many organisms, especially plankton to co-occur in dense swarms or

clumps or aggregates (qv), or nekton, primarily fish, to co-occur in schools or other aggregations.
pathogen

patógeno

An organism that is the causative agent of a disease. Pathogens are usually microbes (viruses, bacteria, fungi, etc.).


pathway

ruta, camino, vía

Possible route or path by which organisms were distributed or spread; synonymous with

corridor (qv) or "track" (qv) (cf barrier, node).
patristic distance

distancia patrística

A measure of the amount of genetically determined change that has occurred between

any two points of a phylogenetic tree (cf cladistic distance, phenetic distance).
pattern

patrón, modelo

Dispersion (qv) or distribution of organisms, materials or phenomena in space and/or time.


PAUP

Phylogenetic Analysis Using Parsimony (PAUP). A widely used package of computer programs for phylogenetic analysis, developed by D. Swofford, Illinois Natural History Survey.


PCA

See principal components analysis.


pedonic

Pertaining to an inhabitant of the bottom community of a freshwater lake.


pelagic

pelágico

(1) In aquatic systems applied to organisms, materials or processes found in the

water column, removed or essentially so, from influence by contact with the bottom or the nearshore.

(2) In ornithology, applied to seabirds that come to land only to breed.


pelagic biogeography

biogeografía pelágica

Study of the distribution of pelagic (qv) organisms incorporating both historical and

ecological approaches to biogeography.
pelagic clupeoid

clupéido pelágico

A member of the fish families Clupeidae or Engraulidae inhabiting pelagic coastal

systems. Certain species are very important in the ecology and fisheries of major

eastern boundary current ecosystems. In many important respects pelagic clupeioids exhibit parallels in their ecology with some important mesopelagic fish groups.


pelagic region

región pelágica

A biogeographically definable subdivision of the global pelagic environment.


pelagium

pelagobios

A sea-surface community of the open ocean (cf neuston).


pelochthium

A mud-bank community.


penetrance

penetrancia

Genetics: The proportion of individuals of a specified genotype who manifest that genotype as

phenotype under a defined set of environmental conditions.
perigean tides

mareas de perigeo

The tides (qv) of increasing amplitude occurring at the time when the moon is nearest the

earth.

periodicity

periodicidad

Events exhibiting cyclicity, recurring either regularly (predictably) or irregularly.


periphyton

perifiton

Organisms attached to or clinging to stems and leaves of plants or other objects

projecting above the bottom sediments of freshwater ecosystems.
perturbation

perturbación

Any disturbance. Physical, chemical or biological in origin, a perturbation produces

measurable change in a community. The permanent effects of perturbation on a

community, if any, are the subject of equilibrium vs nonequilibrium models of community structure.


pessimal

pésimo

Least favorable; used to refer to values of environmental factors that are close to

the tolerance limits of the organism, farthest from the optimal (qv).
pesticide

pesticida

Any material, usually a chemical of human manufacture, that is used against undesirable organisms (generally animals referred to in the vernacular as pests (qv)).


pest

plaga

Any organism, usually an animal, with undesirable characteristics from the human perspective.


- phagous

Suffix; meaning feeding on, eating; eg ichthyophagous, saprophagous, scatophagous, etc.


phaoplankton

epiplancton superficial

The surface plankton (qv) of the upper photic zone, within the top 30 m of the water column.


phenetic distance

distancia fenética

A measure of the difference in phenotype between any two points on a phylogenetic tree

(cf cladistic distance, patristic distance).
phenetics

fenética

Phylogeny: Estimation of relationship by calculation of an overall, unweighted similarity

value; a purely typological (qv) approach to determining and expressing "relationship" (cf cladistics, evolutionary systematics).
phenocopy

fenocopia

An environmentally induced phenotypic variant that resembles the effect of a known

genetically-based variant, eg from gene mutation.
phenocritical period

período fenocrítico

That phase during development at which the expression of a gene is most easily or

visibly affected by externally applied factors.
phenogram

fenograma

A dendrogram (qv) expressing phenetic relationship (unweighted overall similarity) (cf cladogram).


phenology

fenología

The study of the impact of climate on the seasonal occurrence of floral and faunal

elements.
phenon

fenón

A sample or group of phenotypically similar organisms; used in numerical taxonomy to

replace the term "taxon".
phenotype

fenotipo

The totality of expressed characteristics of an individual (whether observed or measured), as a result of interaction between the genotype (qv) and the environment (cf ecophenotypic).


phenotypic plasticity

plasticidad fenotípica

The capacity for marked variation in phenotype as a result of environmental influences

on expression of the genotype during development.
philopatry

filopatria

(1) The tendency of an individual to return to or stay in its home area.

(2) The tendency of members of a stock (qv) or population to return to the natal

breeding/spawning grounds to reproduce as in salmon or Atlantic herring.


- philous

Suffix; meaning loving, thriving in; eg dendrophilous, helophilous, pelagophilus.


phoresy

foresis

Method of dispersal in which an animal clings to the body of a much larger animal of

another species and is carried some distance before releasing its grasp.
photic zone

zona fótica

Zone in which organisms exhibit behavioral or physiological response to day/night changes in light level. (cf euphotic).


photoperiodism

fotoperíodo

The response of an organism to periodic often rhythmic changes in either the intensity

of light, or, more usually, to increasing or decreasing daylength.
photophore

fotóforo

Luminous organ - a discrete morphological structure which contains all of the necessary

chemistry for bioluminescence, normally neuronally controlled - found in a variety of

deepsea (and some coastal marine) fishes and other organisms.


phototrophic

fototrófico

Of or pertaining to organisms that obtain their energy from light reactions; autotrophs (qv) (cf chemotrophs, heterotrophs).


phototropic

fototrópico

Tropic response (directional orientation and/or movement) of an organism to the

stimulus of light; also termed heliotropic (qv) in more direct reference to the sun.
phreatic

freático

Pertaining to ground water.


- phyceae

Suffix; the ending of a name of a class in botanical nomenclature.


- phycidae

Suffix; the ending of the name of a subclass in botanical nomenclature.


phycocoenology

ficocenología

Study of algal communities.


phyletic gradualism

graduaismo filético

New species (forms, series, taxa) arise over time through gradual and continuous

phyletic transformation, believed to proceed at a slow and constant rate; (cf anagenesis).
phylogenetic tree

árbol filogenético

Dendrogram (qv) representing a hypothesis of phylogeny (qv).


phylogeny

filogenia

Evolutionary relationships within and between taxonomic levels, especially the patterns

of lines of descent.
physics

física

Oceanography: Applies to physical oceanography and oceanographers ("physicists"). Essentially the study of the forcing functions (qv) helping to explain dynamic processes in the oceanic hydrosphere and the distribution of oceanic physical properties (temperature, light, pressure, flow fields, etc).


physiognomy

fisionomía

(1) Ecology: The form and structure of natural communities.

(2) Systematics: The body form and appearance of individual organisms.
physiological ecology

ecología fisiológica, ecofisiología

The study of the functioning of organisms in relationship to their environment.


phytobenthos

fitobentos

Autotrophic benthic organisms.


phytogeography

fitogeografía

The biogeography of autotrophs, especially plants. This branch of biogeography is

also known as floristics.
phytoplankton

fitoplancton

Autotrophic (cyanobacteria, protists, plants) plankton (qv).


phytosociology

fitosociología

Description of plant communities, especially their classification based on floristic

rather than life form or other physiognomic criteria.
phytotelmic

fitotélmico

Used of organisms that inhabit small pools of water within or upon plants

(phytotelmata).
picoplankton

picoplancton

Plankton (qv) with maximum dimensions on the order of 0.2 - 2.0 microns


pioneer

pionero

Ecology: The first species or assemblage to colonize or recolonize a barren or disturbed area,

thereby commencing a new ecological succession.
plagio -

Prefix meaning oblique, as in plagiotropism, an orientation response at an

oblique angle to the vertical.
planetic

Motile; possessing motile or swarming stages.


plankter

plancter

An individual planktonic organism; phytoplankter; zooplankter.


plankton

plancton

Pelagic organisms incapable of maintaining their distribution against the movement of

water masses (cf. nekton). Commonly viewed as passive drifters although many are

capable of considerable vertical migration.


plasticity

plasticidad

The capacity of an organism to vary morphologically, physiologically or behaviorally

in response to environmental fluctuations.
plate tectonics

tectónica de placas

Unifying concept encompassing continental drift (qv), seafloor spreading, and other major

dynamic geophysical process including volcanism and seismic events. The concept is one

of a lithosphere of rigid plates of crust and upper mantle material "riding" upon a deformable æsthenosphere. The driving force is presumed to be heat derived through decay of radioactive elements within the earth.


pleio -

Prefix meaning more.


pleiotropy

pleitropía

The phenomenon of a single gene being responsible for a number of different phenotypic

effects.
plenary power

Taxonomy: The authority of the ICZN to suspend the provisions of the Code.


plesiomorphous

plesiomórfico

Phylogeny: Primitive (ancestral) state of a character (as opposed to apomorphous (qv) or

derived)(cf derivative).

pleuston

Organisms permanently found at the sea surface, limited to the surface by their

own buoyancy, often or typically extending into the air, and subject to wind drift (eg Sargassum, Physalia, Velella).
pluvial

(1) Pertaining to or resulting from the action of rain or precipitation.

(2) Used of a geological period or of a climate characterized by abundant rainfall.
POC

COP

Particulate organic carbon. Nonliving detrital suspended material in seawater,

part of the seston (qv), which also includes living particulate matter (cf DOC).
poikilo -

Prefix meaning various, variable.


poikilosmotic

poiquilosmótico

Refers to organisms that do not exhibit regulation of internal salt/fluid content relative

to the external milieu over the range of conditions specified; osmotic conformers.
poikilotherm (ectotherm)

poiquilotérmico

Organism that regulates body temperature by behavioral means only, if at all. In the vernacular termed "cold-blooded" (cf. homoiotherm).


polar

polarOceanic zone, at high latitudes, where sea surface temperatures exhibit an annual excursion from below 0°C to about 5° C.


poly -

Prefix meaning many.


polygamy

poligamia

In animals a pattern of mating in which an individual has more than one sexual

partner (includes polyandry and polygyny).
polygenic character

caracter poligénico

Quantitatively variable character (as expressed phenotypically) which is the result of

interaction of a number of genes.
polyhaline

polihalino

(1) euryhaline (qv).

(2) The second most saline zone of an estuary based on the Venice System (qv) of

classification of brackish waters.


polymictic

polimíctico

Applied to lakes where waters are circulating (overturning) virtually continuously

(eg in high altitudes in the tropics).
polymorphism

polimorfismo

The existence of two or more forms (differing in phenotypic expression) that are

genetically distinct but contained and maintained within the same interbreeding

population.


polynya

polinya

An expanse of open water in the middle of sea ice, often permanent or semipermanent.


polyphyletic

polifilético

Phylogeny: An artificial grouping of taxa based on homoplasy (qv), convergently acquired

apparent synapomorphy (qv), but not true synapomorphy.
polyploidy

poliploidía

Genetics: A condition in which the number of chromosome sets in the nucleus is a multiple (greater than 2) of the haploid numbers.


polythetic

politético

A taxon of whatever rank that is not uniquely diagnosed by one or more autapomorphic (qv)

character states but is defined by a combination of character states, a large proportion of which occur in most of the members but no single feature is uniquely possessed by all of the members.
polytopic

politópico

Occurring in many localities or geographic areas (cf monotopic, syntopic).


polytypic species

especie politípica

Divided into subspecies or genetically distinct populations, varying

geographically (cf monotypic species).
polytypy

politipia

The occurrence of phenotypic variation (cf phenotype) between populations or subgroups within a species that are geographically distinct. The main problem in studying the variation between

such groups is distinguishing between ecophenotypic (qv) vs underlying genetic difference.
pontic

póntico

Pertaining to the deep sea.



pool

existencia

Ecology: The total or partial quantity of a component, compound, material, etc. (cf active pool, reservoir pool).


population

población

An infraspecific subdivision: an assemblage of organisms regarded as members of the

same species, differing from other such assemblages, if any, in relatively panmictic gene

exchange and in local differentiation. Unrigorously defined in most cases, the concept of population lies on the continuum between deme (panmictic) and species (reproductively isolated from other species) (cf stock).


population biology

biología poblacional

The intersect of population ecology (qv) and population genetics (qv).


population ecology

ecología poblacional

The study of populations (qv) in an environmental context, ie the study of physical and biological environmental parameters and their effects on the growth, age-structure and reproduction of populations.


population genetics

genética poblacional

The study of gene frequencies and selection pressures in populations.


population structure

estructura poblacional

The age and sex composition of a population, principally products of survivorship (lx)

and age-specific fecundity (mx).
positive estuary

estuario positivo

A "normal" estuary (qv) whose waters are of lower salinity than the adjacent sea (cf negative estuary, neutral estuary).


post -

Ecology: The total or partial quantity of a component, compound, material, etc. (cf active pool, reservoir pool).

Prefix meaning after, behind, succeeding, later than.
potamodromous

potamódromo

Reproductively migrating from a lake or lentic body of freshwater into a tributary

stream or lotic body of freshwater, eg sea lamprey stocks in the North American Great

Lakes.
potamoplankton



potamoplancton

Planktonic organisms of slow-moving rivers and streams.


potamous

potámico

Pertaining to the lower reaches of rivers and streams


potential niche

nicho potencial

See fundamental niche.


pre -

Ecology: The total or partial quantity of a component, compound, material, etc. (cf active pool, reservoir pool).

Prefix meaning before, in front of, prior to, earlier than.
preadaptation

preadaptación

Adaptation (qv) evolved in one adaptive zone (qv) (habitat, environment) which proves fortuitously advantageous in a different adaptive zone, allowing the organism to radiate into it. No selection for the alternate adaptive zone is implied.


precedence

precedencia

Taxonomy: The order of seniority of available names or nomenclatural acts.


precocial

precoz

Used of offspring or species that develop rapidly in obtaining independent self-maintenance (cf altricial).


predation

predación

Interaction between species populations in which one organism, the predator, obtains

energy (as food) by consuming, usually killing, another organism, the prey. Almost always refers to the consumption of one animal by another (cf grazing).
predator

pradador

The consumer in predation (qv) coaction (qv).


preferential species

especie preferencial

A species that is present in varying abundance in several communities (cf Braun-Blanquet classification), but especially abundant in one particular community (cf accidental, exclusive, indifferent, or selective species).


prey

presa

The consumed in predation (qv) coaction (qv).


primary production

producción primaria

The autotrophic fixation of carbon dioxide by photosynthesis, expressed as a

rate per unit volume or per unitary area of sea surface per unit time. (typical oceanic units:

mg-C/m2/day or g-C/m2/year). Gross primary production is rate of carbon fixation per unit volume per unit time. Net primary production is gross primary production minus respiration, and represents the fraction of captured and converted energy available for population growth of the autotroph and for herbivore consumption.


primary succession

sucesión primaria

Succession initiated on a newly-produced bare area, with no living remnants of a

previously-existing community (if any) (cf secondary succession).
primitive

primitivo

Phylogeny: Preserving the character state(s) of an ancestral stage (taxon). The term may be applied to a single character (as a synonym of plesiomorphic) or to the whole organism or to

a taxon.
primordial

Primitive; original; used of the earliest stage in the ontogeny or development of an

organ or system.
principal component analysis (PCA)

análisis de componentes principales (ACP)

A method of transforming the axes of multidimensional space in which observations occur

such that the first axis explains the maximum amount of variance; the second axis

(orthogonal to the first) explains the maximum of the remaining variance, and so on. The first 2 or 3 axes explain most of the variance.


pro -

Prefix meaning before, in front of, forward.


probability

probabilidad

Statistics: The chance that a given event will occur. The probability of an impossible

event is zero, the probability of an inevitable event is unity (ie 0 <= p <= 1).
production

producción

See primary productivity, secondary productivity.


profundal

de profundidad

Pertaining to the deep zone of a lake, below the level of effective light penetration.


progression rule

regla de la progresión

In cladistic biogeography (qv), the idea that, in general, the most plesiomorphous (qv) members of a monophyletic lineage will be found in that area at or closest to the area of origin of the group, the most apomorphous (qv) will be found in those areas most distant.
prolific

prolífico

Abundant; able to produce large numbers of offspring.


promunturium

A rocky seashore community.


pros (pro) -

Prefix denoting the positive condition.


prospective

potencial, posible, probable

Potential; possible; probable (cf realized).


protandry

protandria

Sequential hermaphroditism in which individuals are initially functional as males and

later switch to being functional females.
protocooperation

protocooperación

An interaction of species populations (qv) in which both populations benefit

but neither is dependent on the relationship (cf mutualism, amensalism, commensalism).
protogyny

protoginia

Sequential hermaphroditism in which individuals are initially functional as females and

later switch to being functional males.
provenance

origen

Biogeography: The place of origin.


province

provincia

See biogeographical province.


provincialism

provincialismo

See endemism.


psamathium

A strandline community of a sandy seashore.



psammon

The microscopic flora and fauna of interstitial species between sand grains.


pseudo -

Prefix meaning false.


pseudo-oceanic

pseudo-oceánico

Species that are members of otherwise pelagic and oceanic groups that are obligatory

or facultative in their strong association with bottom communities. Commonly associated

with continental slopes or island margins.


pseudopelagic

pseudopelágico

Synonym of pseudo-oceanic (qv).


pseudoplankton

pseudoplancton

Organisms not normally planktonic occurring accidentally in the pelagic realm

(cf tychopelagic) .
pseudovicars

pseudovicariantes

"Ecologically equivalent species" (qv); Unrelated or distantly related but ecologically equivalent

species occupying separate and disjunct geographic areas.
pteropod ooze

fango de pterópodos

Calcareous deepsea biogenic fine-grain sediment in which at least 30% of the sediment consists of shells / shell fragments of the small planktonic gastropods known as pteropods.


pulsed forcing

Forcing functions (qv) which cycle off and on (or weaken / intensify) in cyclic or

rhythmic progression, eg the effects on ocean circulation of seasonally occurring monsoonal

wind fields.


punctuated equilibria

equilibrio puntuados

Evolution of new species (forms, varieties, taxa) concentrated in very rapid events,

considered nearly instantaneous in terms of geologic time.
pycnocline

picnoclina

Zone of rapid change of density per unit distance (usually vertically).


pyramids

pirámide

So-called Eltonian pyramids (qv) depict material/energy flow in a community. Commonly cited

transfer rates (producer ==> herbivore ==> carnivore, etc) center on 10%, with the

rest expressed as entropy or transferred to the decomposer chain (cf inverted pyramid).


Q-mode

modo Q

Methods for analyzing data in which the observations (N) (records) form the columns

and the variables or attributes (n) (fields) form the rows in a table or matrix (cf

R-technique).


quadrat

cuadrado

A delimited area for sampling flora or fauna, usually placed randomly, haphazardly or arbitrarily within the study area. A one meter square frame is a typical size. Quadrat may also refer to the

physical sampling frame itself.
quadrature

cuadratura

The time at which the sun and moon are approximately at right angles with respect to

the earth, associated with neap tides (cf syzygy).
qualitative

cualitativo

Descriptive, non-numerical.


quantitative

cuantitativo

Numerical; based on counts, measurements, ratios or other values.


quantum evolution

quantum evolutivo

A "burst" of evolution (cf punctuated equilibria).


quiescent

quiescente

Being quiet, still, or at rest; inactive.


Q10

Ratio of the speed of a reaction at a given temperature to that of the same reaction at

a temperature 10° C lower. This ratio is approximately 2.0 for most biological

reactions.


R
r vs K

Basic terms of the exponential (qv) and (K) logistic (qv) models of population growth.

In ecology a common oversimplification is an attempt to categorize species as either

"r-selected" or "K-selected" (qv).


R-MODE

modo R

Methods for analyzing data in which the observations (N) (records) form the rows

and the variables or attributes (n) (fields) form the columns in a table or matrix

(cf Q-technique). This is the more typical method of data organization.


r-selection

selección r

Selection for maximizing the intrinsic rate of increase of an organism (r), so that

when favorable conditions occur, eg dispersal into a newly opened habitat, the species

is able to expand its numbers rapidly to colonize the area (cf K-selection).


R-strategist

R-estratega

An r-selected species (cf r-selection; C-S-R triangle).


race

raza

Interbreeding group of individuals genetically distinct from the members

of other such groups of the same species. Usually these groups are geographically

isolated (cf allopatry) from one another so that there are barriers to intergroup gene flow.


radiation

radiación

(1) Ecology: any portion of the electromagnetic spectrum impacting life.

(2) Evolution: The evolution of many different forms with many different adaptations

within a single lineage (cf adaptive radiation).


radiolarian ooze

fango de radiolarios

Siliceous deepsea fine-grain sediment in which at least 30% of the sediment consists of tests of the protists known as radiolarians.


rafting

Passive transport of organisms by solid nonliving objects, ranging from rafts

of floating, downed vegetation at the sea surface to transport of entire floras and

faunas via continental drift.


random

al azar, aleatorio

A pattern of distribution where individual organisms are sparse in relationship to the

total area (volume) and in relationship to the total number of samples, and where every

possible sample locality has the same (low) probability of being occupied by the organism.

In other words capture or noncapture in one sample is nonpredictive of capture or noncapture

in a second sample from the same area. This distribution is well-approximated by a Poisson Distribution (cf even, aggregated)


random assemblage

conjunto aleatorio

The concept that a community (qv) of organisms is found together on a random basis

(cf superorganism concept).
random numbers

números aleatorios

Statistics: A table of numbers in which the probability of any number occurring at

any one time is constant and independent of all preceding numbers.
range

área

The particular area occupied by an organism or group of organisms or included within its

(their) ambit (qv).
rank

rango

A product of ordinal scaling (cf scale), the assignment of relative position (first,

second, third, fourth,..., last) irrespective of absolute quantitative difference.
rank-abundance

rango de abundancia

Relative abundance of organisms within a community ordered by rank, with the most

abundant species assigned rank=1, the next most rank=2, and so forth. A common measure of

community similarity in open ocean studies involves comparisons of rank-abundance.


rare

raro

(1) Very seldom occurring; typical sampling distribution fits a Poisson.

(2) Refers to a species known to exist in a community but that is often absent from a series

of samples from that community.


Rassenkreis

circulo de razas

A polytypic species, especially when the populations are naturally arranged in a

zonal or meridional trend line, reflecting gene exchange and/or barriers thereto.
realized

realisado

Actual or observed (cf prospective).


realized niche

nicho realizado, nicho efectivo

Ecology: The actual constraints under which an organism operates - negative coaction such as

competition or predation, suboptimal physical environmental conditions, etc. - ensure

that the niche (qv) in which the organism operates in the real world has less breadth (qv) than that in which it could operate if the only limits were its own physiological tolerances and intraspecific interactive effects (cf fundamental niche).


realm

zona, área

Major biogeographic region, eg Nearctic, Ethiopian (terrestrial); Antarctic, tropical or equatorial (marine). Also used for major ecological regions (eg pelagic, oceanic).


recapitulation

recapitulación

Heterochrony (qv) that results in the appearance during ontogeny of development mirroring presumed rectilinear evolutionary change in a lineage.


recruitment

reclutamiento

Fisheries Biology: First appearance of individuals of a fishable stock in the fishery - younger individuals are not taken due to size or location or both.


rectilinear

rectilíneo

Used of growth or movement that follows a straight line trajectory, also orthogenesis (qv).


recurrent group

grupo recurrente

A group of species that consistently co-occurs in samples from an environment, area or

community. Consistency of co-occurrence is determined by pairwise calculation of

similarity indices using presence/absence data from field samples.


red clay

fango rojo

A pelagic (oceanic) sediment containing less than 30% material of biogenic origin (cf ooze); extremely fine clay mineral particles, accumulating very slowly, typically underlying the most oligotrophic areas of the subtropical anticyclones, covering about 38% of the deep ocean floor.


red tide

marea roja

A marked bloom of aquatic plants, protists or procaryotes; typified by dinoflagellate

blooms discoloring the water a reddish brown coloration, often with concomitant

production of toxins and bioluminescence phenomena. (cf milky seas)


reducers

reducidores

Ecology: Organisms, principally bacteria, that gain nutrition chemolithotrophically in anoxic environments using principally nitrate or sulfate ions as electron receptors, thereby reducing those compounds (eg denitrifying bacteria , sulfate-reducing bacteria).


refugium

refugio

Small isolated area where extensive changes in environmental conditions, most typically

changes in climate, have not occurred. Plants and animals formerly widespread in the

region now find a refuge from the new and unfavorable conditions in such an unaltered location. Alternatively an area or environment in which a species otherwise displaced by competitive exclusion survives.


region

región

See biogeographical region or realm.


regression

regresión

Paleontology/ historical geology: the withdrawl of the sea from a land area (cf transgression).


relationship (phylogenetic)

relación (filogenética)

(1) Evolutionary systematics: the relative closeness of two taxa in an evolutionary sense.

(2) Cladistics: two taxa are each others closest relative (sister taxa) if and only if each

shares with the other a more recent common ancestor than does either with any other

taxon.
relictual distribution

distribución relictual

Applied to the distribution of organisms or taxa, perhaps formerly widespread, now

surviving in a fraction of their former range or in an environment generally more

specialized or less favorable than previously occupied.


remote sensing

monitoreo remoto

Use of aircraft, spacecraft, satellites, and other platforms to detect from afar electromagnetic cues (typically involving sensing of light, heat, uv emissions or reflections) concerning

environmental conditions at the sea or land surface.
reproductive potential

potencial reproductivo

Population biology: the expected average per capita production of offspring per female.


reservoir pool

fondo de reserva

In the biogeochemical cycle, that portion of the nutrient or active substance that is not actively in exchange but may be available to the organisms involved (cf active pool).


residence time

tiempo de residencia

A measure of the chemical or biological reactivity of a substance in seawater and its

rate of removal (typically to the sediments). Residence time is the ratio of the

input/output flux of the material (assumed to be in equilibrium) to the total amount.


resilience

resiliencia

Ecology: A stability metric, the rate at which a community or ecosystem returns to an original state following a perturbation (cf fragility).


resource partitioning


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