328
Shamshev & Grootaert: A review of the genus Stilpon from the Oriental region
2 long bristles in apical part. Epandrium completely divided.
Left epandrial lamella small, fused to hypandrium, lacking
bristles in apical part. Left surstylus with upper lobe (Fig.
38) divided; lower part slender, lacking surstylar comb, upper
part subglobular, with 1 short bristle. Right surstylus (Fig.
39) fairly large, rather elongate oval, with rounded apex,
lacking spines. Left cercus unbranched, slender, short, lacking
spines, with several bristles of different length in basal part.
Right cercus consisting of two lobes, lacking spines; right
lobe hardly prominent, with several marginal bristles of
different length; left lobe short, broad, rounded at apex.
Phallus short.
Female. In most respects identical to male. Terminalia
shortened. Segment 8 normally sclerotized. Proximal margin
of sternite 8 without 2 anteriorly directed rods. Apex of
sternite 8 hinged and partially separated from base. Sternite
10 uniformly sclerotized, not fused with ventroapical margin
of tergite 8. Cercus elongate oval, brownish yellow, clothed
in setulae of different length.
Measurements. – Body length 1.4-1.7 mm, wing length 0.9-
1.2 mm.
Etymology. – “Taksin” (south in Thai) refers to south of
Thailand where this species was found.
Phylogenetic relationships. – The relationships of
S. taksin
are unresolved beyond inclusion within the S. seeluang
species group. However, the presence of 2 very long bristles
on the segment 8 of the male abdomen, the general structure
of the male terminalia and, especially, the greatly reduced
cerci suggest this species is allied with S. laawae, S.
crassinervis, and
S. nhamyaaw.
Distribution and seasonal occurrence. – Thailand. Only
known from one locality in South Thailand. The record is
from October (end of rainy season in South Thailand).
The S. divergens species group
Cumming & Cooper (1992) have recognised this group to
include a single species known just from the Oriental region
only. Additionally, they noted also some undescribed species
from Africa and Japan. We have added to this group another
6 species described below. Members of the S. divergens
species group are characterised by an entirely black thorax
(except S. malayensis) and tomentose scutum, complete rows
of acrostichal setulae, presence of gland-like structures on
the male abdomen (unclear in S. divergens), undivided upper
lobe of left surstylus, markedly developed surstylar comb,
bilobed right cercus, a long ventral bristle on left cercus, setose
left cercus, long bristles in apical part of left epandrial lamella,
elongate hair-like phallus, elongate female terminalia and
sternite 8 of the female terminalia with upper apex not
partially separated from base.
Stilpon khorngkeun, new species
(Figs. 40-44)
Material examined. – Holotype - male,
THAILAND: Loei
province, Na Haeo, FIRS, Malaise trap in bamboo forest at Na Haeo
FIRS, 1 male, 7-14 May.2000, coll. P. Grootaert (SWU).
Paratypes – Same data as in holotype: 1 male, 17-24 Sep.1999;
1 male, 24-30 Apr.2000; 1 male, 13-20 May.2001; 1 male,
29 May.-4 Jun.2001; 1 male, 7-14 Jul.2001, coll. Verapong
Figs. 36-39. Stilpon taksin, new species, male. 36, mid leg, anterior view, 37, hypopygium, ventral view, 38, upper lobe of left surstylus,
dorsal view, 39, right surstylus, dorsal view. Scale bar: 0.1 mm.
329
THE RAFFLES BULLETIN OF ZOOLOGY 2004
Kiatsoonthorn & P. Grootaert; 2 males, Na Haeo FIRS,
Malaise trap n
°
1 in bamboo wood, 25 May.2003 (leg. P.
Grootaert); 3 males, Na Haeo FIRS, base of waterfall, sample
n
°
23029, 16 May.2003 (leg. P. Grootaert). (in coll. SWU,
RBINS & ZRC).
Diagnosis. – Species with black thorax, differing from other
species from the Oriental region by R4+5 strongly curved in
apical part and by brown hind tarsomere 1.
Description. – Male. Head black in ground-colour, with
minute anterior ocellars, short (half as long as inner verticals)
posterior ocellars and long inner verticals. Antenna yellow.
Postpedicel nearly 2.0 times longer than wide. Style about 5
times longer than postpedicel. Palpus brownish yellow.
Thorax brown. Scutum entirely tomentose. Postpronotal
bristle long, inclinate. Dorsocentrals in multiple rows,
complete posteriorly. Acrostichals 2-serial, complete
posteriorly.
Legs with colour pattern: hind femur brownish yellow in
apical 2/3, tarsomere 5 of all legs and hind tarsomere 1 brown;
otherwise legs yellow. Hind trochanter lacking spinules. Mid
femur (Fig. 40) slender, with long pale ventral bristles in
extreme base and 2 ventral yellowish spines nearer to middle.
Hind femur (viewed laterally) evenly thickened toward
middle, with short anterodorsal bristles (3 subapicals longer),
lacking prominent dorsal bristles near base but with prominent
dorsal bristles in apical part. Fore tibia with short, hardly
prominent, brownish, posterodorsal bristle at apex, lacking
prominent ventral bristles. Mid tibia considerably shorter than
mid femur, with rows of ventral spinules and 1 longer
subapical spinule. Hind tibia unmodified.
Wing (Fig. 41) normally developed, covered with uniform
microtrichia, finely infuscate. Costal vein with ordinary short
setulae on anterior margin. Veins R2+3 and R4+5 thickened,
darker than other veins (especially R4+5). Vein R2+3 about
3 times longer than Rs. Distance between apices of R2+3
and R4+5 nearly 1.5 times shorter than distance between
apices of R1 and R2+3. R4+5 strongly curved at apex. R4+5
and M divergent in apical part. Vein CuA1 not quite reaching
wing margin. Halter with brown elongate knob (one specimen
had a very dark knob) and pale stem.
Abdomen largely yellowish brown, bearing mostly scattered
dark setulae which are longer on pregenital segments, with
all tergites (except segment 8) of subequal in length, tergites
1-2 unmodified. Narrow gland-like structures present between
tergites 4-3 and 3-2.
Hypopygium (Fig. 42) brown. Hypandrium with 2 strong
bristles in apical part. Epandrium completely divided. Left
epandrial lamella small, fused to hypandrium, with 2
moderately long bristles in apical part. Left surstylus with
Figs. 40-44. Stilpon khorngkeun, new species, male. 40, mid leg, anterior view, 41, wing, dorsal view, 42, hypopygium, ventral view, 43,
upper lobe of left surstylus, dorsal view, 44, right surstylus, dorsal view. Scale bar: 0.1 mm.